Peer-reviewed journal articles

Clapham M. & Kitchin J. (2016) Social play in wild brown bears of varying age-sex class. Acta Ethologica, 19 (3): 181-188. Link

Clapham M., Nevin O. T., Ramsey A. D. & Rosell F. (2014) Scent marking in wild brown bears: time investment, motor patterns and age-related development. Animal Behaviour, 94: 107-116. Link

Clapham M., Nevin O. T., Ramsey A. D. & Rosell F. (2013) The function of strategic tree selectivity in the chemical signalling of brown bears. Animal Behaviour, 85 (6): 1351-1357. Link

 Clapham M., Nevin O. T., Ramsey A. D. and Rosell F. (2012) A hypothetico-deductive approach to assessing the social function of chemical signalling in a non-territorial solitary carnivore. PLoS ONE 7(4): e35404. Link

Clapham, M., Nevin, O. T. & Darimont, C. T. Assessing the scent marking frequencies of wild brown bears in response to varying population density, food availability and habitat characteristics. In prep for submission to Animal Ecology.

Clapham, M. Darimont, C. T. & Rosell, F. Preliminary chemical analysis of brown bear cutaneous scent. In prep for submission to Journal of Zoology.

Rosell F., Owen M., Clapham M., Tinnesand H. V., Swaisgood R. R., Swenson J. E. & Zedrosser, A. Chemical communication in bears (Ursidae). In prep for submission to Mammal Review.

Please contact me directly with requests for journal articles which are not open-access. 

Presentations at scientific conferences

Rosell F., Clapham M*., Owen M., Swaisgood R.R., Swenson J.E. & Zedrosser, A. (2016) Chemical communication in bears. 24th International Conference on Bear Research and Management, Anchorage, Alaska. *presenting author

Clapham M., Nevin O.T., Ramsay A.D. & Rosell F. (2013) Assessing the scent marking behaviour of wild brown bears; an exploration of time and energy investment, motor patterns and age-related development. 22nd International Conference on Bear Research and Management, Provo, Utah. Download

Clapham M., Nevin O.T., Ramsay A.D. & Rosell F. (2013) Food availability affects the scent marking frequencies of wild brown bears. 22nd International Conference on Bear Research and Management, Provo, Utah. Download

 Clapham M., Nevin O.T., Ramsay A.D. & Rosell F. (2012) Using camera traps to study behaviour in wild populations: a case study of the brown bear Ursus arctos. Invited symposium presentation at the 3rd European Congress on Conservation Biology (ECCB), Glasgow, Scotland. Download

 Clapham M., Nevin O. T., Ramsay A.D. & Rosell F. (2012) Using Markov Chain Analysis to Assess Stereotyped Scent Marking Behaviour in Wild Brown Bears. 14th International Behavioural Ecology Congress (ISBE), Lund, Sweden. Download

 Clapham M., Nevin O. T., Ramsay A.D. & Rosell F. (2011) Inter-sexual and inter-seasonal differences in the chemical signalling strategies of brown bears. 20th International Conference on Bear Research & Management, Ottawa, Canada. Download

 Clapham M., Nevin O. T., Ramsey A. D. & Rosell F. (2010) Preliminary findings assessing the olfactory communication strategies of brown bears. 19^th International Conference on Bear Research and Management, Tbilisi, Georgia. Download

 Clapham M., Nevin O. T., Sinclair B. & Ramsey A. (2009) Connectivity in brown bear populations: an assessment of gene flow in coastal British Columbia. 2^nd European Congress of Conservation Biology (ECCB), Prague, Czech Republic. Download